RESEARCH TEAM
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RESEARCH TEAM
Elena Anagnostopoulou
Principal Investigator
Elena Anagnostopoulou obtained her Ph.D. in Linguistics from the University of
Salzburg in 1994. After a post-doc at MIT (1997-1998), where she returned in
2007 as a Visiting Associate Professor, she took a position at the University of
Crete in 1998, where she is currently Professor of Theoretical Linguistics. Her
research interests lie in theoretical and comparative syntax, formal linguistic
typology, morphology and historical morphosyntax, with special focus on the
interfaces between syntax, morphology, and the lexicon, argument alternations,
Case, Agreement, person, gender, clitics, control and anaphora. In 2013 she
received a Friedrich Wilhelm Bessel Research Award by the Alexander von
Humboldt Foundation, Germany, in recognition of her past accomplishments in
research and teaching. Since 2019 she is elected member of Academia
Europaea. She is the author of The Syntax of Ditransitives. Evidence from Clitics
(Mouton de Gruyter 2003), co-author of External Arguments in Transitivity
Alternations. A Layering Approach (Oxford University Press 2015), has co-edited
several collective volumes. She has published in journals (Natural Language and
Linguistic Theory, Linguistic Inquiry, Language, Syntax, The Journal of
Comparative Germanic Linguistics, among others) as well as in edited volumes
and conference proceedings. She is Co-editor in the Series Open Generative
Syntax. Language Science Press and member of the editorial board of the
journals Journal of Greek Linguistics, Linguistic Inquiry and Syntax. She was
international Co-Investigator in the AHRC-funded project "Investigating variation
and change: Case in Diachrony" (PI: Christina Sevdali) at the University of Ulster
(2017-2019). She is PI of the UCrete-funded project "Using tools from
Evolutionary Biology to reveal relationships among languages” (2019-2023) in
collaboration with Manolis Ladoukakis (Department of Biology) and of the
H.F.R.I- funded project “Modeling Glossogeny.” (2019-2022; in collaboration with
Manolis Ladoukakis and Dimitris Michelioudakis, AUTH).
Manolis Ladoukakis
Research team member
Manolis Ladoukakis received his Bachelor in Biology from the Aristotle University
of Thessaloniki, Department of Biology and his M.Sc. in Marine Biology from the
Department of Biology, University of Crete, Greece. He received his Ph.D. from
the same Department. He worked as a Marie Curie Post-doctoral fellow at the
Centre for the Study of Evolution, University of Sussex, UK, and as a visiting
Lecturer at the School of Life Sciences, University of Sussex, UK. He was elected
in 2007 as a Lecturer at the Department of Biology, University of Crete, Greece,
where he is currently an Associate Professor. In 2015 he worked at the
Department of Ecology and Evolutionary Biology, Brown University, RI, USA as a
Fulbright fellow. He teaches the “Evolution” and the “Molecular Evolution”
courses at the Department of Biology, and he participates in other undergraduate
and postgraduate courses. His research interests regard the evolution of the
inheritance and recombination of mitochondrial DNA and the co-evolution of
nuclear and mitochondrial genomes. He is also interested in the factors that
determine the structure of microbial communities. Further, he works on the
extension of the evolutionary theory in other evolvable systems beyond
organisms, such as microbial communities (in collaboration with ecologists) and
languages (in collaboration with linguists). His research has been published in
several journals such as Molecular Biology and Evolution, PLoS Biology, Trends
in Ecology and Evolution, Molecular Ecology and others. He has participated in
several research projects. He is a collaborator in the UCrete-funded project
"Using tools from Evolutionary Biology to reveal relationships among languages”
(2019-2023) with Elena Anagnostopoulou (Department of Philology, Division of
Linguistics) and in the H.F.R.I- funded project “Modeling Glossogeny.” (2019-
2022; in collaboration with Elena Anagnostopoulou (PI) and Dimitris
Michelioudakis, AUTH).
Dimitris Michelioudakis
Research team member
Dimitris Michelioudakis studied classics and linguistics at the University of Athens
(BA 2005) and theoretical linguistics at the University of Cambridge (MPhil 2007,
PhD 2012). He was teaching and research assistant at Cambridge (2010-2013),
where he taught introductory and intermediate courses in theoretical linguistics,
syntax, semantics and typology, and postdoctoral research associate at the
University of York (2013-2018), where he taught intermediate and advanced
courses in comparative and diachronic syntax, the syntax-semantics interface
and syntactic typology. Since 2019, he is an Assistant Professor of General
Linguistics at the Aristotle University of Thessaloniki. He specialises in the syntax
of Greek and its dialects, syntactic change, comparative syntax and its interfaces
with semantics (mainly argument and event structure), pragmatics and
morphology (compounding and case). His research interests also include syntax-
based phylogenetics and the use of syntactic parameters to probe genetic
relatedness and the intensity of language contact. In 2015, he conducted
fieldwork on two South American languages in Brazil, as part of his British
Council-funded Researcher Links project. He has participated in international
conferences and published a number of book chapters in edited volumes,
conference proceedings, and articles in journals such as Natural Language and
Linguistic Theory, The Linguistic Review, Rivista di Grammatica Generativa, and
Glossa. Since December 2019, he is part of the research team of the HFRI-
funded project “Modeling Glossogeny” alongside Elena Anagnostopoulou (PI,
Department of Philology, Division of Linguistics, University of Crete) and Manolis
Ladoukakis (Department of Biology, University of Crete).
Maria-Margarita Makri
Postdoctoral Researcher
Maria-Margarita Makri has a degree in Philology with specialisation in linguistics
from the University of Athens (2011) and completed her postgraduate studies at the
University of Cambridge (MPhil 2011) and the University of York (MA by Research
2014, PhD 2019) with funding from the ESRC and the A.G.Leventis Foundation. She
has taught undergraduate courses in linguistics at the department of Language and
Linguistic Science of the University of York, the department of Pedagogy of the
University of the Aegean and the postgraduate programmes of the School of
Humanities of the latter. In 2019 she joined the Department of Mediterranean
Studies at the University of the Aegean as a postdoctoral researcher and her project
“Degree constructions in Standard Modern Greek and its Dialects” is funded by the
Greek State Scholarships Foundation. In March 2021 she joined as a postdoctoral
researcher the ModelGloss project at the University of Crete, which is funded by the
HFRI. Her research interests lie in comparative syntax, semantics, language
acquisition and experimental linguistics. She has presented her research in
international conferences and seminars (GLOW, NELS, SuB, GALA, LAGB, MIT
LingLunch, MIT LFRG, Cambridge Syntax Lab a.o.) and published in international
journals (Glossa, Rivista di Grammatica Generativa) and conference proceedings.
Pavlos Pavlidis
External Collaborator
Pavlos Pavlidis has received his PhD from the University of Munich in the lab of
Prof. Wolfgang Stephan. He worked on population genomics of natural populations of
Drosophila, disentangling adaptation from stochastic non-adaptive processes such as
random genetic drift. Following graduation, he worked with Prof. Alexandros
Stamatakis at HITS, Heidelberg, developing and optimizing algorithms for whole-
genome analysis.
His current research interests are related to population genomics and
bioinformatics. He is interested in the evolution of gene regulatory networks by
means of drift and positive selection, the co-evolution between different loci, the
spatial coalescent and the connections between evolution and the 3D structure of
DNA. Furthermore, he is conducting research on the analysis of microarray data
focusing on gene expression regulation.
Athanasios Michael Ramadanidis
Collaborating Researcher
Athanasios Michail Ramadanidis is a PhD candidate in Theoretical Linguistics at the
University of Crete since 2018. He studied Linguistics and Romance Studies at the
University of Vienna, where he received his Master's degree in 2017. His research
interests focus on theoretical syntax, morphology, compositional semantics, and
their interface, adpositions, Case, and spatial expressions. His doctoral research "On
the Hierarchy of Case and Adpositions in Synchrony and Diachrony" is funded by the
Hellenic Foundation for Research and Innovation (HFRI, 2019–2022) and is
supervised by Prof. Elena Anagnostopoulou (principal advisor), Prof. Winfried
Lechner (National and Kapodistrian University of Athens), and Prof. Susanne
Wurmbrand (University of Vienna / Harvard). Since 2019 he has been collaborating
with Prof. Elena Anagnostopoulou and Assoc. Prof. Manolis Ladoukakis on the
research project "Using tools from Evolutionary Biology to reveal relationships
among languages" (2019–2023) which is funded by the University of Crete and as a
volunteer external collaborator on the project "Modelling Glossogeny" funded by
HFRI (2019–2022). Athanasios Michail Ramadanidis is also a musician and a graduate
in Piano Performance from the Department of Music Science and Art of the
University of Macedonia.
Christos Zioutis
External Collaborator
Christos Zioutis studied Biology at the University of Crete (2009-2013). He has been granted a scholarship and graduated from the graduate programme “Molecular Biology and Biomedicine”, jointly ran by the Departments of Biology and Medicine, University of Crete and The Institute of Molecular Biology and Biotechnology (IMBB) of the Foundation for Research and Technology Hellas (FORTH) (2013-2015). In his graduate studies, he worked on the inference of positive selection and demography in the human gut microbiome, under the supervision of Dr. Pavlos Pavlidis. He is currently a PhD candidate (2015-2022) at the Centre for Microbiology and Environmental Systems Science, at the University of Vienna, supervised by Univ- Prof. Dr. David Berry and his PhD research is focused on evolution in ecologically meaningful timescales, by studying the polysaccharide degraders of the human gut microbiota. His work is a combination of experimental evolution in germ-free mice and NGS technologies. His main scientific interest is the evolution and dynamics of biological systems - mainly of microorganisms and microbial communities.
Ruby Sleeman
Research Assistant
Ruby Sleeman is a PhD candidate in General Linguistics at Goethe University Frankfurt, Germany, since 2017. Her doctoral research “The syntax of ordinals” in the project “Nominal Modification”, funded by the German national research foundation DFG, focuses on the differences in morphosyntax between Dutch and German expressions containing ordinal numerals and superlative adjectives (“the second-highest mountain”). Her main research interests include morphosyntax (minimalism/generative syntax), specifically of the nominal domain; numeral expressions, language variation and change, corpus linguistics, and the interaction between syntax and semantics. She obtained her Research MA in Linguistics from Leiden University, the Netherlands (2017). During her MA studies in Leiden, she was a teaching assistant and additionally worked as a research assistant on the project “Language and Number” in Amsterdam, building a crosslinguistic typological database of morphosyntactic variation in numeral expressions (funded by the Dutch national research foundation NWO). In April 2022 Ruby Sleeman joined the interdisciplinary research project “Modeling Glossogeny” funded by the HFRI as a Research Assistant. She has presented her research in various international conferences and workshops (SLE, SinFonIJA, IGG, ATINER, DeMiNeS Masterclass, a.o.)